Invertebrates are animals without vertebrae. This group includes arachnids (spiders & scorpions), crustaceans, gastropods (snails, slugs, etc), insects, and others. While an estimated 97% of all animals are invertebrates, their tracks and sign are often overlooked by trackers. Insect tracks are among the most encountered of all invertebrate tracks and they are an extremely diverse and interesting group of animals to explore.

Leaf-cutterAnt Mounds
Leaf-cutterAnt Mounds
Ant Tracks
Ant Tracks
Ant Tracks
Ant Trail
Antlion
Antlion Trail
Antlion Traps
Banana Slug Slime
Beetle and Peromyscus Tracks
Beetle Eating Grasshopper
Beetle Tracks
Beetle Tracks
Beetle Tracks
Caterpillar Scat (Frass)
Caterpillar Scat (Frass)
Caterpillar Tracks
Caterpillar Tracks
Caterpillar Tracks
Centipede Tracks
Centipede Tracks
Chinese Mantis Ootheca
Cicada Holes
Cicada Holes
Cicada Exuvia
Cicada Exuvia
Crab Tracks (Peru)
Crab Tracks (Peru)
Crayfish and Raccoon Tracks
Crayfish Burrow
Crayfish Chimney
Crayfish Tracks
Crayfish Tracks
Crayfish Tracks
Cricket Tracks
Cricket Tracks
Dragonfly Larva Tracks
Dragonfly Track
Dung Beetle
Dung Beetles
Ghost Crab Feeding Sign
Ghost Crab Feeding Sign
Ghost Crab Feeding Sign
Ghost Crab Tracks
Ghost Crab Tracks
Hermit Crab Tracks
Roundworm Tracks
Jerusalem Cricket Tracks
Jerusalem Cricket Tracks
Jerusalem Cricket Tracks
Jerusalem Cricket Tracks
Jerusalem Cricket Tracks
Jerusalem Cricket Tracks
Katydid Scat (Frass)
Katydid Scat (Frass)
Leafcutter Bee Sign
Leafcutter Bee Sign
Millipede Tracks
Millipede Tracks
Mole Cricket Ridges
Mole Cricket Ridges
Mole Cricket Ridges
Mole Cricket Ridges
Mole Cricket Ridges
Mud Dauber Digs
Mud Dauber Digs
Mud Dauber Digs
Mud Dauber Digs
Mussel Predation by Raccons
Praying Mantis Ootheca
Say's Stink Bug
Scorpion Tracks
Scorpion Tracks
Scorpion Tracks
Scorpion Tracks
Scorpion Tracks
Scorpion Tracks
Slug Feeding Sign
Slug Feeding Sign
Slug Scat
Snail Scat
Sow Bug Tracks
Sphinx Moth Larva Scat (Frass)
Spider Burrow
Spider Burrow
Spider Leaf Nest
Spider Tracks
Spider Tracks
Swallow Digs and Mud Dauber Digs
Tarantula Tracks
Tarantula Tracks
Twig Girdler Sign
Twig Girdler Sign
Twig Girdler Sign
Twig Girdler Sign
Walking Stick Tracks
Walking Stick Tracks
Wasp and Burrow
Worm Scat (Castings)
Worm Scat (Castings)
Worm Scat (Castings)
Worm Scat (Castings)
Worm Scat (Castings)
Worm Scat (Castings)
Worm Trail
Insect & Invertebrate Tracks Gallery Full Species List
- Ant Mounds
- Ant Tracks
- Ant Trail
- Antlion Trail
- Antlion Traps
- Banana Slug Scat
- Banana Slug Slime
- Beetle Eating Grasshopper
- Beetle Tracks
- Caterpillar Frass
- Caterpillar Scat
- Caterpillar Tracks
- Centipede Tracks
- Chinese Mantis Ootheca
- Cicada Holes
- Cicada Shells
- Crab Tracks
- Crayfish Burrow
- Crayfish Chimney
- Crayfish Tracks
- Cricket Tracks
- Dragonfly Larva Tracks
- Dragonfly Track
- Dung Beetle Scat
- Ghost Crab Feeding Sign
- Ghost Crab Tracks
- Hermit Crab Tracks
- Jerusalem Cricket Tracks
- Katydid Scat
- Leafcutter Bee Sign
- Millipede Tracks
- Mole Cricket Ridges
- Mud Dauber Digs
- Mussel Predation by Raccons
- Praying Mantis Ootheca
- Roundworm Tracks
- Say’s Stink Bug
- Scorpion Tracks
- Slug Feeding Sign
- Slug Scat
- Snail Scat
- Sow Bug Tracks
- Sphinx Moth Larva Frass
- Spider Burrow
- Spider Leaf Nest
- Spider Tracks
- Swallow Digs and Mud Dauber Digs
- Tarantula Tracks
- Twig Girdler Sign
- Walking Stick Tracks
- Wasp and Burrow
- Worm ScatCastings
- Worm Trail
Arachnids (spiders and scorpions) have 8 legs and therefore leave tracks with groups of 4 dots on each side of the trail. Sometimes it can be quite difficult to make out all 4 dots when the hindmost foot disrupts the marks from a previous foot.
Crustaceans (primarily crayfish and crabs here) have quite distinctive tracks. Crayfish walk primarily on 8 legs and have small claws at the end of their legs. If you look closely at their tracks you will find pairs of dots from each part of the claw. This is a very helpful characteristic for identification. Also look for two drag marks created by the sides of the tail. Crabs have 8 legs and walk sideways and leave a repetitive somewhat tire-tread like trail. Some species of crabs like horseshoe and hermit crabs leave distinct patterns that allow for easy identification.
Gastropods (mainly slugs and snails here) leave a number of signs including: slime trails, scats, and feeding signs. Slugs often feed on algae on tree bark and other surfaces and leave distinctive patterns in the process. Snail and slug scat forms long twisted ropes that are easily identified in the field.
Insects are an enormous group of animals. Insects that often leave tracks include: ants, beetles, crickets, grasshoppers, mantids, millipedes, wasps, and many, many more. Insects can be very difficult to identify from an actual specimen and identifying them to the species from tracks can at times be impossible. Sometimes the best we can do is identify to the order level.
The following images are the recent invertebrate track observations from the North American Animal Tracks Database. This is an iNaturalist project where trackers share observations and help each other learn about animal tracks, all while contributing to scientific research.
mediumOchre Sea Star
I think these don't really like crossing open sand, so don't leave tracks very often.Cecropia Moth
mediumVertebrates
mediumVertebrates
Not sure what passed this pure fur
Alligator or raccoonVertebrates
mediumVertebrates
medium
mediumAnimals
Small tracks in snowVertebrates
mediumGray Sanddragon
tracksBark and Ambrosia Beetles
Bark beetle galleries in logEarthworms and Allies
tracksGray Sanddragon
tracksCeanothus Silk Moth
cocoonEarthworms and Allies
Tracks and probe hole from a spotted sandpiper in earthworm casting. This observation for the earthworm casting.Spiders
Strands of gossamer webbingEarthworms and Allies
Muddy trail on a rock next to riverArthropods
tracksEarthworms and Allies
tracksEarthworms and Allies
mediumGray Sanddragon
tracksSpiders
mediumMillipedes
scat under logGlover's Silkmoth
mediumVertebrates
Tracks in snow, seems likely to be a catArthropods
tracksOysters and Allies
Shells from an old pile of shells that got bulldozed.Arthropods
tracks
Despite the fact that invertebrate tracks and sign are all around us and frequently encountered, there are very few guides available that will help you identify their tracks. Aside from the comprehensive coverage of the topic in Tracks and Sign of Insects and Other Invertebrates, most field guides barely scratch the surface. I’ve included a few of the guides that include insects or invertebrates. If you decide to purchase any of these guides, using the links below will help support future developments to this website.
Tracks & Sign of Insects and Other Invertebrates opened my eyes to an entire new world of animal tracking. Charley Eiseman and Noah Charney included an incredible amount of insect sign, from feeding signs to egg cases to scats. This guide is a valuable resource for anyone interested in learning about insects and it is a great way for trackers to expand their knowledge. These guys also have some great videos on identifying insect tracks and sign on their website.
Animal Tracks and Scat of California is a regional guide, but many of the species are found throughout North America. This guide includes mammals, birds, and even some reptiles, amphibians, and invertebrates. It is a great general tracking guide (full disclosure: I am a co-author of this guide) and is a great bet if you want a single guide that covers more than just mammals. This book includes some drawings of various insect and invertebrate trails.
Wildlife of the Pacific Northwest is a regional guide written by my friend and excellent tracker David Moskowitz. While this book primarily focuses on the Northwest, a lot of information applies nationwide. This tracking guide includes mammals, birds, reptiles, amphibians, and invertebrates. It dedicates 8 pages specifically to insects and other invertebrates at the end and has a really cool section on identifying beetle galleries created in tree bark. The guide also contains excellent original artwork by Dave and some incredible photography.